I really like it, I think that’s great. A few tweaks:
At the moment all the oceans are adjacent, if it’s like this then I guess all the life in the columns will be very similar as they will be quite well mixed. I think it would be interesting to break the middle left connection so if you want to go from ocean o1 to ocean o2 you have to go through a shallow sea s1. Imagine a band of land all round the planet maybe. That way there’s two large oceans connected so we can see what happens there and one which is a bit seperated.
I think it might interesting to have an ocean column of a shallower depth, maybe a small arctic ocean? It would also give an area which is almost always frozen which would be interesting I think, one of the vents could be moved there.
Here it is with those changes, what do you think? Otherwise it’s great.